The Western Highland
Rim of Tennessee is prominent on aerial photos or satellite images as being
overwhelmingly forested. Within this region of the state, the Swan Creek
watershed stands out for having a number of rare plants and invertebrates,
as well as being connected to some key pieces of public land.
While some
parts of the region have been converted to pine plantations, it is mostly
allowed to regenerate following harvest of the hardwood timber. Aside from
scattered pieces of state and federal public lands, including Wildlife Management
Areas and State Parks, it is overwhelmingly private land, Parts are in small
holdings and parts in larger tracts owned by forest industry.
The Tennessee Natural
Heritage Program has been aware of the significance of the biological
diversity of the Swan Creek watershed for several years, and has conducted
several inventories on the area. The Heritage Program, part of the Ecological
Services Division of the Tennessee Department of Environment and Conservation
(TDEC/ESD), maintains a database of the locations of Tennessee's rare
plants and animals. The information in this database is used to identify
lands in need of conservation, and is consulted by numerous public agencies
who use it in planning projects in order to avoid impacts to natural resources
and biodiversity.
A Natural Areas Registry
provides a way to afford recognition and provide management recommendations
to landowners and public agencies whose land contains habitat for rare
plants and animals. In recent years, additional
populations of several rare plants have been located in the Swan Creek
watershed.
The most globally rare of these is the Federal Endangered Tennessee
yellow-eyed-grass, Xyris tennesseensis. This small plant with grass-like
leaves is not really a grass, but is a member of a small specialized family
the Xyridaceae. Members of the genus Xyris have yellow flowers which emerge
from a cone-like structure at the top of a leafless stem.
The Tennessee yellow-eyed-grass
is found in Tennessee, Alabama, and Georgia; one of the largest known
populations is in the watershed of Big Swan and Little Swan Creeks. This
species in found only in a specialized habitat called a "seep-fen" where
cold spring water flows across a shallow soil composed of limestone-like
gravel mixed with organic muck derived from the breakdown of plant material.
One of the five Tennessee sites is on the lands of the Natchez Trace Parkway
and is Registered as a State Natural Area with the Tennessee Natural Heritage
Program.Others are on private land; one landowner agreed to restrict
logging in the vicinity of the plants due to efforts of the Tennessee
Field Office of the Nature Conservancy. More populations need to be located
and protected for this species' survival to be assured.
Several other plants
which are listed as rare in Tennessee are found in this "seep-fen" habitat.
There are similar habitats in Missouri and New York. The other state rare
plants include the Large-leaved grass-of-Parnassus (Parnassia grandifolia),
the Fen orchis (Liparis loeselii), and the Small-headed rush (Juncus brachycephalus).
These are all plants which are more common in other parts of the United
States, but which are rare in Tennessee.
The Grass-of-Parnassus is not
a grass at all, but a plant with beautiful five-petaled white flowers
and kidney-shaped leaves; the flowers open in the late fall, when not
many plants are blooming. This seep-fen habitat
is particularly rare in Tennessee, and occurs only in small patches which
are isolated from one another. The physical characteristics of the small
seeps provide the special conditions which the rare plants require in
order to thrive. Frequently, if the seeps are large enough, there is not
enough soil for trees to be rooted and grow. These open seeps receive
more sunlight, and herbs or shrubs dominate the ground layer.
The Tennessee
yellow-eyed grass is most often found in this type of site. The Grass-of-Parnassus
can grow in even smaller seeps which are under the shade of forest trees.
Some sites of this type are found on the upper reaches of Swan Creek;
often native lilies occur here as well, but they do not bloom reliably
in the shade.
Another interesting feature of limestone seeps is that they
may provide habitat for dragonflies. The adults lay their eggs in the
loose wet soil of the seeps. When the eggs hatch, the larvae are able
to live in the clean, clear water which seeps out of the sloping surfaces.
The sunny areas around the seeps provide areas where the adults can forage
for prey during the summer. A new species of dragonfly, the Tennessee
Snaketail (Ophiogomphus acuminatus) has recently been described from Lewis
County, Tennessee.
One current project of collaboration between Tennessee
Natural Heritage and TVA Entomologists is to determine if this rare dragonfly
shares the same habitat as any of the rare pants of the Lewis County seeps
or if any rare plants can be found a the sites where ihe dragonfly occurs.
Rare plant conservation
is most valuable when efforts to conserve species result in better understandings
of the functions and values of the systems which contain them. Rare species
frequently serve as the visible indicators of systems which we may not
fully understand.
Another globally rare
plant which lives in a very different habitat is Eggert's sunflower (Helianthus
eggertii). Yellow sunflowers frequently confound the identification efforts
of all but the most determined botanists. Actually, this species is relatively
easy to identify if one knows exactly what to look for. It blooms in July
and August, like most sunflowers; its flowers (actually composite heads
of many small flowers) are relatively large (about 3.5 inches across),
its stem is smooth and waxy, and the tapering leaves with rounded bases
are smooth except for a scattered roughness on the upper surface. This plant is globally
quite rare, and has been a Federal Category 2 (C2) Candidate for a number
of years.The U.S. Fish and Wildlife Service is close to listing it as
Threatened. It prefers a habitat type which was presumably more widespread
when fire was a more common event in the landscape.
This grass and herb-dominated
habitat type is called "barrens", and is related to the prairies of the
Midwest, both in structure, species composition, and ecology. Presumably, when fire
occurred more frequently, and large grazing animals (such as bison) roamed
free, there were large areas of parts of Tennessee and the Southeast which
had relatively few trees, with abundant stands of native grasses and flowering
herbs, like composites and legumes. Under present conditions, this community
persists on roadsides and recently disturbed areas.
In Tennessee, Eggert's
sunflower is most frequent in the Lewis/Lawrence County area and in Coffee
County on the Eastern Highland Rim. A large population is known from the
lands of the Arnold Engineering Development Center (AEDC) near Tullahoma.
Several stands are known from roadsides or clearings on the land of The
Farm, near Summertown, and from nearby highway rights-of-way.
As the highway
sites are vulnerable to destruction during road widening, the management
of sites on large tracts of public land and on the lands of sympathetic
private landowners is critical to the species' survival.
The conservation of
lands in the Swan Creek and Little Swan Creek watershed is important not
only as part of a largely forested portion of the state which benefits
many species, including neotropical migrant songbirds but some specific
habitat types provide homes for several rare plants. In particular, the
globally rare Tennessee yellow-eyed-grass and Eggerts sunflower have significant
populations in the Swan Creek watersheds.
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